Naran, Daksha, Skelton, Paul H, Villet, Martin H. Karyology of three evolutionarily hexaploid southern African species of yellowfish, Labeobarbus Rüppel, 1836 (Cyprinidae). 2007.
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Naran, Daksha, Skelton, Paul H, Villet, Martin H. African Zoology; 2006. Karyology of the redfin minnows, genus Pseudobarbus Smith, 1841 (Teleostei: Cyprinidae): one of the evolutionarily tetraploid lineages of South African barbines.
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Marriott, Michael S, Booth, Anthony J, Skelton, Paul H. Reproductive and feeding biology of the Natal mountain catfish, Amphilius natalensis (Siluriformes: Amphiliidae). 1997.
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Skelton, Paul H, Cambray, James A, Lombard A, Benn, G. Patterns of distribution and conservation status of freshwater fishes in South Africa. 1995.
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Skelton, Paul H. Fifty years of ichthyology in Grahamstown. 1996.